I feel like the fire should be animated. And faster.
Also, the top of the tower feels pretty crowded. Might just be under intense assault, but this seems to play into the hands of the defenders.
The central fire feature in the tower is extraneous. After four or five runs through the chamber, the outer left and right path (past the fast moving spike block) is far easier (and faster) to navigate. The water feature feels really well thought out, though it feels less than intuitive at the moment. Having the water triggered by co-op chain pulling is nice, but it might be cool to not have the 'timer' that causes the flood to end on it's own, forcing defenders to interact with the chains to set up the traps in the main room again.
If this is the way towering is going, I'm excited to see what comes.